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A giant ray appears jubilant as it smiles widely while being fed. The diver had entered the tank to check for any dead fish - but the ray spotted an opportunity for a snack.

When they ran out of food the rays were still reluctant to let the diver go, and followed her around as she tried to leave. The photos were taken at the Enoshima Aquarium in Kanagawa, Japan, by professional British photographer David Williams, who said: "I was originally going to take photos of...
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